The story is told in eight major parts, called Scenes.

Scene One begins in 1846, at Combe-Raven in West Somerset, the country residence of the wealthy Vanstone family: Andrew Vanstone, his wife, and their two daughters. Norah, age 26, is happy and quiet; Magdalen, 18, is beautiful but volatile and willful. They live in peace and contentment, looked after by their governess, Miss Garth.

Through amateur theatricals, Magdalen discovers she is a talented actress and falls in love with Frank Clare, the idle but handsome son of a neighbour, who is also in the play. They want to be married, and their fathers agree. Although Frank fails at every career he reluctantly tries, and his father is not wealthy, Magdalen's fortune will easily support the young couple.

But before they marry, Mr. Vanstone is killed in a train crash and Mrs. Vanstone dies in childbirth. The girls discover from the lawyer, Mr. Pendril, that their parents have only been married for a few months, and their wedding invalidated the will which left everything to the daughters.

Since the daughters are illegitimate, they have no name, no rights, and no property. Combe-Raven and the entire family fortune are inherited by Andrew's older brother, Michael Vanstone, who has been bitterly estranged from the family for many years. He refuses to provide any support for the orphaned young women. With the help only of their governess Miss Garth, they set out to make their own way in the world.